https://phys.org/news/2018-07-berry-gorging-disperse-seeds-scat-small.html

In southeastern Alaska, brown and black bears are plentiful because of salmon. Bears frequently supplement their salmon-based diet with fruit as they build their fat stores for winter hibernation. As a result, their seed-filled scats are found throughout the landscape.

"Salmon can have a far greater impact on the ecosystem than we thought," said study lead author Yasaman Shakeri, an Oregon State University graduate now with the Alaska Department of Fish and Game. "Our study shows how small mammals can benefit indirectly from salmon through high bear densities that salmon support and the resulting seed-filled scats on the landscape. Not only are small mammals spending months feeding and fighting for the seeds in scats, they're also scattering the seeds on the landscape, which allows some of the seeds to become future fruiting plants."